1 Introduction

The Yangtze River and its catchment occupy one of the largest drainage-basin systems in eastern Asian continent and act as a boundary zone of northern and southern China (e.g. Zhao and Chen 1999; Wu et al. 2012). The lower reaches of the Yangtze River (LYR) are situated in an area that is dominated by the East Asian monsoonal circulation (EAM); and the natural vegetation is warm temperate broadleaf and subtropical evergreen forests as well as transitions of these biomes (e.g. Ren and Beug 2002; Innes et al. 2014). The vegetation and environment in the LYR region would have been sensitive to even small-scale Holocene climatic as well as sea level fluctuations, due largely to its special bioclimatic locality and any marked weakening or strengthening of the EAM regime (e.g. Morrill et al. 2003; Yi et al. 2003; Zong et al. 2011). In particular, the LYR has been identified as one of the birthplaces for prehistorical Chinese civilization such as the ancient Hemudu and Liangzhu Cultures, and for rice cultivation such as the domesticated paddy Oryza sativa, as broadly elucidated by detailed palaeoecological and archaeological evidence obtained in earlier studies (e.g. Londo et al. 2006; Zong et al. 2007; Atahan et al. 2008; Fuller et al. 2009; Zhao et al. 2009; Li et al. 2010). As a consequence, the LYR has thus been suggested as an ideal region of interest for exploring the history of Holocene environmental, climatic, and cultural changes in eastern monsoonal China (e.g. Chen et al. 2005, 2009; Innes et al. 2009, 2014; Wang et al. 2011, 2012; Zong et al. 2007, 2011, 2012).

Quantitatively-integrated and high-resolution records of Holocene precipitation and temperature variations are rarely available in the LYR region, and the nature of Holocene climatic evolution in this region remains unclear (e.g. An 2000; Yi et al. 2003; Chen et al. 2005; Zong et al. 2006, 2011, 2012; Atahan et al. 2008; Innes et al. 2009; Shu et al. 2010). Several questions arise, including (1) has the Holocene climate evolved similarly or differently between the LYR and other EAM-affected regions of China such as northern monsoonal China? (2) What potential driving factors have possibly triggered the Holocene climatic oscillations in the LYR? (3) Have precipitation and temperature patterns behaved synchronously or asynchronously during the course of Holocene in the LYR? (4) What was the timing and magnitude of the Holocene climatic optimum in the LYR? Making progress on these issues requires regional-scale, high-resolution, and numerical palaeoclimatic data.

Quantitative palaeoclimatological estimates using biological fossil proxies preserved in sediments provide an important avenue to develop these climatic data for comparison with regionally-averaged and temporally-detailed Holocene precipitation as well as temperature data in northern China (e.g. Xu et al. 2010; Chen et al. 2015; Li et al. 2015, 2017), the Tibetan Plateau (e.g. Shen et al. 2008; Herzschuh et al. 2009; Wang et al. 2014a, b), and many regions of Europe and North America (e.g. Seppä et al. 2009; Bartlein et al. 2011; Viau et al. 2012; Heiri et al. 2014; Mauri et al. 2015; Ladd et al. 2015), at local, regional, and continental scales. The continuous and conceptual advances in developing and improving large-scale calibration datasets and novel statistical techniques, have greatly promoted the accuracy and robustness of climatic reconstructions, with different means of transforming a variety of fossil biological assemblages into numerical estimates of past rainfall and temperature during the Holocene (e.g. Birks et al. 2010; Salonen et al. 2014; Wang et al. 2014a, b; Li et al. 2015, 2017). Such reconstructed climatic data are particularly valuable for validating climatic transient-model simulations and other independent climatic proxies comprising various geophysical, geochemical, and geoecological records from both terrestrial and marine environments worldwide (e.g. Braconnot et al. 2012; Renssen et al. 2012; Heiri et al. 2014; Li et al. 2015, 2017). In this respect, pollen data are one of the most commonly-used biological proxies for quantitative Holocene terrestrial climatic inferences at a broad scale (e.g. Seppä et al. 2009; Birks et al. 2010; Wang et al. 2014a, b; Li et al. 2017).

In view of the above, we place a key focus on reporting new, high-resolution Holocene precipitation and temperature reconstructions based on three fossil pollen sequences located within the lower Yangtze region of monsoonal China. Our main purpose is to enable a broader understanding and discussion with respect to the Holocene climate-related issues presented above regarding the LYR and monsoonal China. We further assess our pollen-based Holocene climatic reconstructions by relying upon an extensive multi-proxy comparison with a large number of either single or integrated moisture- and temperature-related proxy records from China as well as other regions of the world, such as speleothem oxygen isotope records (e.g. Fleitmann et al. 2003; Wang et al. 2005), loess-palaeosol sequences (e.g. Wang et al. 2014a, b; Li et al. 2014a, b), lake sediment cores (e.g. Shen et al. 2005; Chen et al. 2015), and climate model simulations (e.g. Jin et al. 2013; Chen et al. 2014). Moreover, the most important element of this study is to contribute improved and meaningful insights for evaluating the role of causal forces and atmospheric circulations in driving Holocene climatic, environmental, and cultural changes as well as on forecasting their future possible dynamics in the lower Yangtze region of China.

2 Study area

The lower valley of the Yangtze basin is located in the eastern part of monsoonal China (Fig. 1), where the Asian monsoon dominates the climate and is characterized by warm, wet monsoon in summertime and cold, dry monsoon in wintertime, with four well-pronounced climatic seasons occurring per year. The 700 hPa atmospheric airstream lines in Fig. 1 clearly show that the lower Yangtze region is under the typical influence of the East Asian summer monsoon system, whilst northwestern and southwestern China are dominated by the westerlies and Indian summer monsoon circulation, respectively. Mean annual temperature (TANN) varies between 14.5 and 16.2 °C, and mean total annual precipitation (PANN) ranges from 800 to 1400 mm, with its maximum level taking place in July (e.g. Yi et al. 2006; Li et al. 2012). The area of our investigation is naturally covered by a high-density of water-bodies including lakes, peats, bogs, marshes, swamps and wetlands, and it has thus been particularly attractive for palaeoenvironmental studies (e.g. Tao et al. 2006; Innes et al. 2009, 2014; Wang et al. 2011, 2012). The vegetation of the LYR area is a biogeographical ecotone consisting of mixed broad leaved deciduous and evergreen forests with distinct transitional characteristics (e.g. Wu 1983; Huang and Zhang 2000). The main temperate components comprise species of Betula, Ulmus, Alnus, Populus, Quercus, and Acer. The dominant evergreen components include Castanopsis, Cyclobalanopsis, Lithocarpus, and Fagaceae. In addition, a low number of coniferous plant taxa such as Pinus, Picea, and Cupressaceae occur in the high-elevation mountainous areas (e.g. Zong et al. 2011, 2012). It is therefore likely that in the transitional vegetation belt of the LYR, the Holocene climatic variability would have caused a northward or southward shift of the temperate or subtropical biome because of fluctuations in precipitation as well as temperature controlled primarily by the overall variability of Eastern Asian summer monsoonal (EASM) intensity (e.g. Yi et al. 2006; Zong et al. 2007, 2011).

Fig. 1
figure 1

Mean 700 hPa atmospheric airstream lines between June and August based on the NCEP/NCAR Reanalysis data during the time interval of 1971–2000 (Kalnay et al. 1996). Blue plus signs correspond to the localities of three fossil sites (Chaohu, Gucheng, and Pingwang) in the lower reaches of the Yangtze basin. ISM, EASM, and Westerlies indicate the regions dominated by the Indian and Eastern Asian summer monsoon, as well as westerly air circulation, respectively. Green dashed line shows the present-day northern boundary of the Asian summer monsoon system modified from Chen et al. (2008)

3 Materials and methods

Pollen-based quantitative estimates for both PANN and TANN were prepared from three Holocene pollen datasets at Chaohu (Chen et al. 2009), Gucheng (Yang et al. 1996), and Pingwang (Innes et al. 2014), that lie within the lower reaches of the Yangtze catchment (Fig. 1). These fossil pollen profiles were selected as they have reliable chronological control and fine-scale temporal resolution (see Table 1 for other details). The AMS radiocarbon technique was utilized to create geochronological datasets (Yang et al. 1996; Chen et al. 2009; Innes et al. 2014). Plant macrofossils, shell fragments, pollen residues, charcoal particles, and basal peats have been used for AMS dating (Yang et al. 1996; Chen et al. 2009; Innes et al. 2014). The age–depth models were estimated by a linear interpolation between adjacent samples (Yang et al. 1996; Chen et al. 2009; Innes et al. 2014). The radiocarbon dates were calibrated and transformed to calendar years before present according to the IntCal13 calibration dataset (Reimer et al. 2013). The AMS 14C dates are presented here as cal. yr BP throughout the text. The three fossil sites are geographically very close to each other, suggesting that they have probably witnessed similar climatic histories during the Holocene.

Table 1 Summary of fossil pollen datasets for PANN and TANN reconstructions at Chaohu, Gucheng and Pingwang in the lower Yangtze region of China

Pollen-based numerical calibration models for both PANN and TANN were established using the Chinese surface pollen–climate database which has been shown to be robust and reliable for Holocene climatic inferences that have been described elsewhere in more detail (e.g. Zheng et al. 2008, 2014; Li et al. 2014a, b, 2015, 2017). The technique in terms of weighted averaging-partial least squares (WA-PLS; ter Braak and Juggins 1993) regression and calibration was chosen for constructing the pollen-based reconstruction models for PANN as well as TANN, because it has been successfully employed in a large number of empirical, theoretical and practical studies; and has been shown to perform as well as or better in comparison with other statistical regression approaches that are commonly applied for developing pollen-based calibration models for regional, continental and global scales (e.g. Birks 1998; Seppä et al. 2009; Birks et al. 2010; Salonen et al. 2012; Li et al. 2015, 2017). The performance of all WA-PLS models was assessed with the method of leave-one-out cross-validation (Birks 1998). The calculated model statistics for PANN and TANN embrace coefficient of determination (R2) between measured and predicted data, root-mean-square-error of prediction (RMSEP), and maximum bias. The two-component WA-PLS models (Fig. 2) were selected with respect to high R2, low RMSEP and maximum bias, as well as the smallest number of useful components (Birks 1998). All terrestrial pollen taxa were taken into account and their percentage values were square-root transformed to reduce noises and stabilize variances in the pollen data (Prentice 1980). The constructions or evaluations of all WA-PLS models associated with the numerical PANN or TANN estimates were carried out using the C2 software (Juggins 2007). In addition, it has been demonstrated elsewhere that PANN and TANN are not always strongly correlated and that either PANN or TANN is statistically significant as well as ecologically meaningful in influencing broad-scale pollen distribution, indicating that they can be employed simultaneously for the pollen-based palaeoclimatic reconstructions applied here (e.g. Li et al. 2014a, b, 2015).

Fig. 2
figure 2

Numerical performance of the calibration models for mean total annual precipitation (PANN) as well as mean annual temperature (TANN) based on pollen data. Statistical parameters consist of coefficient of determination (R2) between observed and predicted values, root-mean-square-error of prediction (RMSEP), and maximum (Max.) bias

To obtain the underlying characteristics and summarize the potential regional signals, a total of six quantitative climatic reconstructions based on the three fossil pollen datasets were integrated to produce two general high-resolution Holocene climatic sequences (PANN and TANN) for the lower Yangtze region of China. Such a composite methodology has been successfully used for quantitative pollen-derived and regional-scale Holocene climatic reconstructions in Europe (e.g. Seppä et al. 2009; Salonen et al. 2014; Mauri et al. 2015), North America (e.g. Viau et al. 2006, 2012; Ladd et al. 2015), and northern China (e.g. Li et al. 2015, 2017). It can be described briefly as follows. Each PANN or TANN estimate was calculated as deviations from their mean values across the Holocene. All estimated values of individual reconstructions for PANN or TANN were then combined so as to prepare two Holocene climatic records with an average time-resolution of circa 36 years for both PANN and TANN. Power spectra for potential periodicities in the pollen-stacked precipitation and temperature records were performed using the Redfit software (Schulz and Mudelsee 2002). This software is able to deal with unevenly distributed time-series data, and can test statistical significance of the spectral peaks against the red-noise background with a null-hypothesis, which can be evaluated by utilizing the first-order autoregressive signals, where characteristic time-scales as well as sampled time-spans match those of the real data, without the necessity for any interpolation (Schulz and Mudelsee 2002).

4 Results and discussion

4.1 Climatic reconstructions with low- and high-frequency trends

The pollen-inferred site-specific Holocene climatic reconstructions are presented in Fig. 3. The regionally-averaged estimates are shown in Fig. 4. Thus we provide the first ~36-year resolution pairwise-dataset for both PANN and TANN records covering the last 10,000 years. These portray both general patterns as well as detailed features of the Holocene rainfall and temperature variations for the lower reaches of the Yangtze catchment. Power spectrum analyses conducted for the composite reconstructions reveal significant periodicities of 4000, 1190, and 116 years for PANN, and of 2500, 1190, and 116 years for TANN (Fig. 5). Some of these periodicities can also be found in other Holocene Asian monsoonal records and various solar parameters (e.g. Li et al. 1996; Laskar et al. 2004; Dykoski et al. 2005; Wang et al. 2008; Wanner et al. 2008, 2011; Zhao and Feng 2014), suggesting a possible mechanistic connection among these climatic systems.

Fig. 3
figure 3

Pollen-based quantitative reconstructions for PANN with RMSEP of 232.06 mm and TANN with RMSEP of 3.62 °C at Chaohu, Gucheng and Pingwang over the last 10,000 years

Fig. 4
figure 4

Pollen-based PANN and TANN estimates during the last 10,000 years for the lower Yangtze region of China derived from the six reconstructions in Fig. 3

Fig. 5
figure 5

Power spectrum analyses for the pollen-stacked PANN and TANN records spanning the past 10,000 years

Overall high-frequency PANN and TANN fluctuations concurrently show a maximum level between 10,000 and 7000 cal. year BP, and a generally declining pattern with strong oscillations from 7000 cal. year BP to the present-day (Fig. 4). The early to middle Holocene had high values of PANN and TANN, suggesting a warm, wet climate and a strong East Asian summer monsoon, as also demonstrated by rapidly increased sea level and possible small paddy rice cultivation in the LYR coastal plain (e.g. Innes et al. 2009; Shu et al. 2010; Zong et al. 2011; Wang et al. 2012). Subtropical forests were regarded as the dominant regional biome during this time phase, with Castanopsis and Cyclobalanopsis identified as the major tree taxa, which is evident in many published fossil pollen records from different parts of the LYR region (e.g. Chen et al. 2005; Yi et al. 2006; Innes et al. 2009; Wu et al. 2010; Li et al. 2012). However, since approximately 7000–5000 cal. year BP, the cool temperate forest elements such as Quercus, Betula and Alnus started to occur and expanded while subtropical elements decreased, eventually leading to a mixed temperate–evergreen vegetation type which prevailed until modern times. The relative sea level was stable with only small fluctuations observed for this time period (e.g. Huang and Zhang 2000; Shu et al. 2007; Yi et al. 2006; Innes et al. 2014). This is closely in line with our climatic reconstructions displaying a falling trend for PANN and TANN values during this period, implying a cooling and drying climate associated with a gradually weakening EASM intensity from the middle to late Holocene in the LYR area (Fig. 4). In addition, it is noteworthy that in eastern monsoonal China, human activity played an important role in influencing vegetation cover during the late Holocene especially the last 2000 years (e.g. Zhao et al. 2009), which implies that pollen-based climatic inferences should thus be treated with caution (e.g. Li et al. 2014a, b).

Several low-frequency cooling and drying episodes can be clearly detected from our pollen-estimated PANN and TANN records. These cold and dry climatic events at about 5300, 4200, 2800 cal. year BP and during the Little Ice Age interval in the LYR region (Fig. 4) appear to be correlated well with those widely reported from other areas in China (e.g. Wang et al. 2005; Shao et al. 2006; Xu et al. 2010; Innes et al. 2014), the North Atlantic region (e.g. Bond et al. 2001; Seppä et al. 2009), and the Northern Hemisphere (e.g. Clemens 2005; Viau et al. 2006; Wanner et al. 2008, 2011). Such short-lived climatic events with high amplitude have been suggested to play a significant role in driving the demise and termination of Neolithic civilization such as the Liangzhu Culture in the Yangtze lowland area, because prehistorical farming fields and human settlements were usually situated close to water bodies such as lake shores and river channels, leading to these agricultural lands and cultural systems becoming vulnerable to abrupt changes in water supplies and thermal conditions during the Holocene, for example during the Neoglacial epoch (e.g. Zhang et al. 2005; Yao et al. 2008; Huang et al. 2010; Wu et al. 2010; Innes et al. 2014).

4.2 Comparison with other subtropical or tropical Holocene records and possible mechanisms

A decadal- to centennial-scale comparison of our pollen-based PANN and TANN estimates for the LYR with other related Holocene climatic proxy records from subtropical China, the Indian summer monsoon (ISM)-influenced region and other tropical or subtropical regions of the world, indicates a macro-scale in-phase pattern of general variability, exhibiting a consistent early Holocene climatic optimum in precipitation or temperature but a relatively dryer or cooler middle to late Holocene (Fig. 6). However, it is notable that this general pattern is different to that indicated by two Holocene hydrological records from the middle reaches of the Yangtze River, that is, the mass accumulation rates of hopanoids from Dajiuhu peat bog as a proxy for water level, and the ratio of anhysteretic remanent magnetization (ARM) to saturation isothermal remanent magnetization (SIRM) from a stalagmite in Heshang Cave as a magnetic proxy, both suggesting a dry middle Holocene but a wetter early or late Holocene (Xie et al. 2013). These two records are also inconsistent with other proxy-based climatic records from the same region, for example, the pollen-based TANN record from Dajiuhu peat (Zhu et al. 2008) and the stalagmite δ18O record from Heshang Cave (Hu et al. 2008), which are in good agreement with our Holocene PANN and TANN records for the LYR. This discrepancy may result from the differences in the various reconstruction techniques, distinct temporal resolutions, reliability of sedimentary chronologies, climatic significance of diverse proxy indicators, or spatial representativeness of sampled sites (e.g. Liu et al. 2015; Rao et al. 2016; Chen et al. 2016a). In addition, the aforementioned Holocene climatic records with consistent patterns that cover a broad geographical region include the following proxy datasets: the mean summer solar insolation (Laskar et al. 2004); stalagmite δ18O records from Dongge (Dykoski et al. 2005), Sanbao (Wang et al. 2008) and Qunf (Fleitmann et al. 2003) Caves; a model-simulated PANN record for southwestern China (Chen et al. 2014); a pollen-inferred PANN estimate for Xingyun Lake in the ISM-influenced Yunan Province (Chen et al. 2014); a TANN reconstruction from Dajiuhu wetland in the EASM-dominated Hubei Province (Zhu et al. 2008); pollen-estimated moisture index (Zhao et al. 2009) as well as tree cover reconstruction (Tian et al. 2016) for southern China; a humification record from Hongyuan wetland on the southeastern Tibetan Plateau (Yu et al. 2006); a synthesized moisture index based on various proxy data for the southern Tibetan Plateau (Ran and Feng 2013); total organic carbon (Yancheva et al. 2007) and tree pollen records (Wang et al. 2007) from Huguangyan Maar Lake in southern China; an ISM rainfall index based on 92 monsoonal moisture records in eastern Asia (Wang et al. 2010); the Intertropical Convergence Zone (ITCZ) index inferred from Ti contents in Cariaco Basin, Venezuela (Haug et al. 2001); a Globigerina bulloides record from a marine sediment core in Arabian Sea (Gupta et al. 2005); composite sea surface temperature (SST) records for Western Pacific Warm Pool (WPWP; Stott et al. 2004; Koutavas and Joanides 2012); and an El Niño/Southern Oscillation (ENSO) record from Laguna Pallcacocha, Ecuador (Moy et al. 2002). Of these records, it is worth noting that the Holocene stalagmite δ18O data have been lately argued to probably represent a signal of the isotopic composition of precipitation from the ISM-influenced region rather than the EASM-dominated territory (e.g. Chen et al. 2014, 2016a; Yang et al. 2014; Wang et al. 2014a, b; Liu et al. 2015).

Fig. 6
figure 6

Holocene comparison of pollen-estimated a PANN and b TANN sequences for the lower Yangtze region (LYR) of China with other related climatic records including: c mean summer solar insolation (SI) at 65°N (Laskar et al. 2004); stalagmite δ18O series from d Dongge (Dykoski et al. 2005), e Sanbao (Wang et al. 2008), and f Qunf (Fleitmann et al. 2003) Caves; g Kiel Climate Model (KCM)-based PANN simulation for southwestern China (Chen et al. 2014); pollen-derived h PANN reconstruction for Xingyun Lake in Yunan Province (Chen et al. 2014) and i TANN reconstruction for Dajiuhu wetland in Hubei Province (Zhu et al. 2008); pollen-composited j moisture index (MI; Zhao et al. 2009) and k tree cover (TC; Tian et al. 2016) records for southern China (SC); l humification (H) record from Hongyuan wetland on the southeastern Tibetan Plateau (Yu et al. 2006); m moisture index (MI) synthesized from various proxy data from the southern Tibetan Plateau (ST; Ran and Feng 2013); records of n total organic carbon (TOC; Yancheva et al. 2007) and o tree pollen percentage (TP; Wang et al. 2007) from Huguangyan Maar Lake in SC; p Indian summer monsoon (ISM) index synthesized for monsoonal China (Wang et al. 2010); q Intertropical Convergence Zone (ITCZ) index from Venezuela (Haug et al. 2001); r Globigerina bulloides percentage record from ODP Site 723 in Arabian Sea (AS; Gupta et al. 2005); s, t composite sea surface temperature (SST-1 and SST-2) records for Western Pacific Warm Pool (WPWP; Stott et al. 2004; Koutavas and Joanides 2012); and u El Niño/Southern Oscillation (ENSO) index from Ecuador (Moy et al. 2002). The grey band depicts the moisture or thermal maximum during the early Holocene in southern China

The above multiple lines of evidence suggest a strong causal linkage between tropical and subtropical climatic systems from the perspective of hemispheric or global scale teleconnections (Fig. 6). It has been often proposed in earlier studies that the orbitally-controlled variability of summer solar insolation would have essentially modulated and triggered the tropical and subtropical summer monsoonal evolution during the post-glacial Holocene epoch (e.g. An 2000; Gupta et al. 2005; Fleitmann et al. 2003; Wang et al. 2008; Zhao et al. 2009). During the early Holocene, the high output of summer insolation may have caused the northward migration of both mean ITCZ and Western Pacific Subtropical High (WPSH) positions as indicated clearly by the high SST values in the WPWP and the Arabian Sea. This therefore resulted in a great amount of evaporated water vapor being transported from the tropical and subtropical oceanic areas to southern and eastern Asia, leading to the northward expansion and penetration of the overall summer monsoonal strength as well as its extensive rain band, and thus abundant rainfall as well as high temperature in the low-latitude regions of monsoonal Asia (e.g. Stott et al. 2004; Zhao et al. 2007; Zhou et al. 2009; Sun and Feng 2013). In contrast, during the middle to late Holocene, a reduced summer insolation would have brought about the southward retreat of both ITCZ and WPSH zones along with the decreased pattern of tropical SST values, which led to a lower moisture transportation from the ocean to continental lands, thereby causing the southward shift and weakening of the summer monsoonal intensity and its rain belt, and reduced rainfall as well as lower heat content in the southern part of monsoonal Asia and China (e.g. Dykoski et al. 2005; Wang et al. 2008; Sun and Feng 2013). In addition, an overall stepwise intensification of ENSO activity from the early to late Holocene (Fig. 6) has been suggested to bring warm water masses with higher SST to the eastern Tropical Pacific Ocean, resulting in lower SST in the western Tropical Pacific Ocean, thus progressively reducing the transport of water vapor to monsoonal China during the entire Holocene (e.g. Wang et al. 2000; Higginson et al. 2004; Sun and Feng 2013).

4.3 Comparison with Holocene climatic records from northern monsoonal and the westerlies’ region of China

A detailed comparison of our inferred precipitation and temperature reconstructions for the LYR with other high-resolution Holocene moisture and temperature records in the monsoonal regions of northern China and in the westerly-dominated areas of northwestern China, reveal an out-of-phase pattern of overall climatic shifts and a major offset in relation to timings of the Holocene climatic optimum interval (Fig. 7). These early published records consist of the following datasets: pollen-reconstructed annual rainfall series from Gonghai Lake (Chen et al. 2015) on the Chinese Loess Plateau (CLP) and Daihai Lake (Xu et al. 2010) in central Inner Mongolia; a multi-proxy based temperature reconstruction for northern China (Hou and Fang 2011); an EASM index based on various proxies in the northern part of monsoonal China (Wang et al. 2010); tree pollen percentages from Qinghai Lake (Shen et al. 2005) and Bayanchagan Lake (Jiang et al. 2006) in northern monsoonal China; a pollen-based moisture record for the northern EASM marginal region (Wang and Feng 2013); frequencies of palaeosol occurrences in the CLP (Wang et al. 2014a, b) and northern China (Li et al. 2014a, b); frequency in the formation of loess or aeolian sands in northern China (Li et al. 2014a, b); a magnetic susceptibility record from the Yulin loess–palaeosol section on the northern CLP (Lu et al. 2013); and a moisture index synthesized from different proxy records for the westerly-influenced regions of Arid Central Asia (ACA) including northwestern China (Chen et al. 2008).

Fig. 7
figure 7

Holocene comparison of pollen-inferred a PANN and b TANN records for the lower Yangtze region (LYR) of China with other climatic proxy records from northern China and other regions of the northern Hemisphere: pollen-reconstructed PANN records for c Gonghai Lake (Chen et al. 2015) on the Chinese Loess Plateau (CLP) and d Daihai Lake (Xu et al. 2010) in northern China (NC); e multi-proxy-based TANN sequence for NC (Hou and Fang 2011); f East Asian summer monsoon (EASM) index for monsoonal China (Wang et al. 2010); tree pollen records from g Qinghai Lake (Shen et al. 2005) and h Bayanchagan Lake (Jiang et al. 2006) in NC; i pollen-based moisture index (MI) for the EASM marginal region (Wang and Feng 2013); frequency distributions of palaeosol occurrences in j CLP (Wang et al. 2014a, b) and k NC (Li et al. 2014a, b); l frequency formations of loess or aeolian sands (AS) in NC (Li et al. 2014a, b); m magnetic susceptibility (MS) record from the Yulin loess–palaeosol section on CLP (Lu et al. 2013); n moisture index (MI) synthesized for Arid Central Asia (ACA; Chen et al. 2008); melt water input (MI) from o the Laurentide Ice Sheet (Jennings et al. 2015) and p the Agassiz Ice Cap (Fisher et al. 2012); q ice-sheet coverage in northern Hemisphere (NH; Dyke 2004); and r Atlantic meridional overturning circulation (AMOC) index based on mean sortable silt grain size (SSGS) from core GS06-144 08GC in northern Atlantic Ocean (Mjell et al. 2015). The grey band shows the moisture or thermal maximum during the middle Holocene in northern monsoonal China. The pink band indicates the moisture or thermal maximum during the early Holocene in southern monsoonal China

The multiple Holocene climatic records, spanning a large geographical area in both northern and northwestern China, point to an overwhelming signal with respect to a middle Holocene climatic optimum that is characterized by the highest precipitation amounts or the warmest temperature conditions (Fig. 7). Recently, Chen et al. (2016b) indicated that in the Xinjiang region as a core area of ACA, magnetic susceptibility records from four loess–palaeosol profiles have exhibited an increasing moisture pattern from the early to late Holocene. However, these trends are out-of-phase with a typical signal of an early Holocene climatic optimum demonstrated by our pollen-stacked PANN or TANN reconstructions for the LYR region as well as other Holocene monsoonal records from the subtropical and tropical domains of China or the ISM-dominated regions presented in this study (Fig. 6). Such an inconsistency may have arisen from unreliable dating controls, various proxy interpretations and resolutions, or different methodological issues and assumptions (e.g. Zhao et al. 2009; Cai et al. 2010; Sun and Feng 2013; Liu et al. 2015). In addition, the climatological viewpoints with regard to this notable timing mismatch remain under debate, which can be tentatively attributed to the significant cooling effect at a hemispherical-scale caused by the deglaciation of broad-scale remnant ice sheets in the high-latitude territories of the Northern Hemisphere during the early Holocene (Fig. 7) (e.g. Chen et al. 2008; Renssen et al. 2012; Li et al. 2015; Liu et al. 2015; Mjell et al. 2015). The rapid melting of these remnant ice sheets such as the Laurentide Ice Sheet (Jennings et al. 2015) in North America and the Agassiz Ice Cap (Fisher et al. 2012) in Greenland, would have likely yielded a large body of freshwater discharge into the northern Atlantic Ocean (Fig. 7). This may have resulted in a suppressed Atlantic meridional overturning circulation (AMOC) as well as North Atlantic Deep Water circulation (NADW), and at the same time increased Ice-Rafted Debris (IRD) in ocean sediments, massive glacial advances, strong meridional temperature gradient and westerly airstreams, and intensified Siberian High and winter monsoon strength, which may in turn have played an important role in blocking the northward movements of the summer monsoon as well as its rainfall front to the northern part of monsoonal China during the early Holocene (Fig. 7) (e.g. Zhao et al. 2009; Chen et al. 2015; Li et al. 2015; Liu et al. 2015).

5 Conclusions

Here we present a pollen-based Holocene climatic dataset with a ~36-year resolution for both PANN and TANN records over the last 10,000 years for the lower reaches of the Yangtze basin in eastern monsoonal China. Our reconstructions show that precipitation and temperature have a concurrent trend of variability on a centennial- to multidecadal timescale, implying a notable climatic pattern of moist–warm or dry–cool intervals during the Holocene. Multi-proxy comparisons indicate that regional-scale Holocene rainfall and thermal variations in the lower Yangtze area are in good agreement with other Holocene climatic records from southern monsoonal China or the ISM-dominated regions, suggesting an early Holocene climatic optimum that was characterized by high precipitation and warm conditions, and a drying or cooling climate for the middle to late Holocene. The orbitally-triggered changes of summer solar insolation and tropical or subtropical climatic circulations such as ITCZ, WPSH, and ENSO may be recognized as the important driving factors for the Holocene summer monsoon variability in the lower Yangtze region of China. Further regional inter-comparisons reveal that the LYR Holocene climatic development has been different to the Holocene moisture or thermal records from the EASM-influenced northern China and the westerly-affected northwestern China where the Holocene climatic optimum mostly took place during the middle Holocene. Overall, our pollen-based high-resolution climatic dataset may be useful for validating climate model simulations, understanding the nature of monsoon climate, and predicting future climatic scenarios in monsoonal Asia and its surrounding areas. Clearly, more case studies would enhance understanding the nature of Holocene climatic changes in different bioclimatic regions of monsoonal China.